Per Ola Karis
Associate Professor, lecturer
karis@botan.su.seBotaniska institutionen
Stockholms universitet
Lilla Frescativägen 5
SE-106 91 Stockholm
Sweden
Phone +46 (0)8 163768
Fax +46 (0)8 165525
My main research interest is the large flowering plant family Asteraceae. I have extensive experience of
anatomy and morphology throughout the family owing to painstaking studies of thousands of representative species
from all major clades. I also work with molecular techniques. Since a couple of years I have returned to my
research "home" South Africa, and I am involved in the joint project "Diversification and evolution in time and
space of Asteraceae in arid and semi-arid southern Africa". By using the target clades Arctotideae, Senecioneae-Othonninae,
and Calenduleae we aim at elucidating species-level relationships in the arid and semi-arid areas, identify
supposed refugia for the Asteraceae (and thus other plant groups) during past climates, model possible range
changes under future climates (e.g. global warming).
In a recent paper based on morphological data I show that
subtribe Gorteriinae (tribe Arctotideae) comprise two sister groups (also supported by recent molecular results
by Vicki Funk et al.) in which both arid area and fynbos radiations has taken place. My PhD student, Frida Stångberg,
works on the Gorteria clade, where the genus Hirpicium seems to be paraphyletic. Frida will also work on the Metalasia clade,
here examplified by Metalasia tricolor on the Rooiberg (Little Karoo), a rare species that was re-discovered
some ten years ago by my friend and colleague, the eminent Jan Vlok (of Oudtshoorn).

The Gorteriinae display an extraordinary array of anatomical and morphological features, and one unique
character are the longitudinally striate hairs that diagnose the Gorteria clade. There is phylogenetically
informative information in the endothecium of the anthers of the Asteraceae, and the Gorteriinae is no exception.
Below are SEM pictures of endothecial cells from a sunflower (Helianthus, right) and from a Berkheya (left).
Note the striations on the perpendicular ribs in the sunflower. One of my aims is to investigate the endothecia
of the Gorteriinae by means of SEM in order to get a clear three-dimensional picture of these diverse cells.
I have also worked on the family Plumbaginaceae, and was involved in an Italian project on the Caryophyllaceae (in particular the genus Moehringia), in which my colleague and friend Simone Fior recently got his PhD at the University of Milan.
Links
The International Compositae Alliance homepage
South African collaborators at Rhodes University, Grahamstown
CapeNature, the Conservation authority of the Western Cape Province
South African National Parks
ALUKA, material about Africa, for example, search images of herbarium sheets of African plants
Publications since 2001
Korall, P & Karis, P. O. 2001. On the typification of Selaginella nova-hollandiae (Selaginellaceae). Taxon 50: 841-844.